Of those I would keep, I think fear has to be top of the list. Although it does sometimes get in the way, it still protects us from taking excessive risks, all be it business or social decisions rather than whether to hunt that vicious squirrel..

Evolutionarily speaking at individual gene lines level, health, fertility, desire to reproduce and traits which help attracting a mate still seem to be key factors (whos got that sense of humour gene…).

On the higher “survival of the species” level, it’s harder to say, but I’d guess it’s likely to be very much social qualities. Collective risk-adversion might help us not to destroy one other on purpose, while long term planning, teamwork and behaving in the interests of the herd rather than the individual may stop us doing it by mistake; not sure how you design for that though, over to you…